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Organising Committee:
Henrik Enghoff, Nils Møller Andersen, Niels Peder Kristensen,
Nikolaj Scharff
Secretariat:
Vibeke Boye, Jytte Regenburg
Assistants:
Marie Grangaard, Niels Grøngaard, Line Sørensen
Lectures and posters are presented at the August Krogh Institute,
Universitetsparken 13,
DK-2100 Copenhagen Ø.
Lunches are available on 29-30 July and 1-2 August at the H.C.
Ørsteds Institute,
upon presentation of lunch tickets. See the map.
Official congress address:
Zoologisk Museum, Universitetsparken 15, DK-2100 København
Ø, DANMARK
Telephone +45 35 32 11 03 (direct) / +45 35 32 10 00
Telefax +45 35 32 10 10. E-mail HEnghoff@ZMUC.KU.DK
The Tenth International Congress of Myriapodology, like its
predecessors, presents lectures and posters on a wide variety
of myriapodological themes. This broadmindedness in approach to
and methodology in the study of myriapods and onychophorans is
a major strength of our congresses: Systematists may be stimulated
by hearing about advances in ecological or ultrastructural research.
Ecologists and physiologists may be inspired to view their results
in the light of new hypothesis on myriapod evolution etc. etc.
While structuring the scientific program it appeared that the
invited and registered lectures could be roughly grouped according
to eight main themes which might be given more or less fanciful
names, as follows:
Ancestry and relationships - Phylogenetic and evolutionary aspects
Shultz & Regier's invited lecture opens this theme, and the entire scientific programme of the congress. Jeffrey W. Shultz will introduce myriapodologists to the technique of molecular phylogeny and its application to myriapods. A lecture on the myriapods' relations to other arthropods, two on centipede evolution, and one on fossil millipedes, are also included under this theme.
Being an arthropod - Segmentational aspects
Arthropods are segmented animals, and no arthropods are more segmented than myriapods. It is therefore appropriate that myriapodologists pay attention to the strong development in research on the genetic basis of segmentation which is taking place in these years. Several lectures are devoted to this subject, including the invited paper presented by Michaelis Averof. A single lecture on postembryonic development is also included here, this subdiscipline of myriapodology being so intimately connected with the segmentation theme.
Order in the diversity - Systematic aspects
After the Rio conference in 1992, the word biodiversity is on everybody's lips. It is the task of systematists to try to organize knowledge of biodiversity or, in our case, "myriapododiversity". Six lectures treat systematic aspects among millipedes and centipedes.
Myriapods on Earth - Geographical aspects
Myriapods, especially millipedes, are of particular interest to biogeographers because of their very poor abilities of dispersal. Sergei I. Golovatch will illustrate this in an invited lecture on Eurasian millipedes. As a counterpart, Luis A. Pereira in another invited lecture will treat the South American fauna of geophilomorph centipedes. Four further lectures are included in this category which, admittedly, is only vaguely delimited vis-a-vis the following one.
Myriapods in their environments - Ecological aspects
This is obviously the most popular aspect of myriapodology, if one can take the number of registered lectures as any kind of indication. No less than 14 speakers will deal with a wide spectrum of myriapod ecology, most emphaszing the myriapods themselves but a few focusing on other organisms (beetles and microbes) which interact with myriapods in Nature. It is particularly welcome that a considerable number of the ecological papers concern tropical myriapods, the emphasis having previously been mostly on European ones.
"A centipede was happy quite" - Behavioural aspects
A centipede was happy quite
until a toad in fun
said, "Pray which leg moves after which?"
This raised her doubts to such a pitch,
she fell exhausted in a ditch,
not knowing how to run
Locomotion has been an important subject in studies of myriapod behaviour. This congress features two lectures on centipede locomotion, and one further on millipede behaviour.
Myriapods magnified - (Ultra)structural aspects
To fully understand our myriapods, we must know what they look like, at all scales of magnification. Three lectures focus on minute details of myriapod morphology, with functional aspects being touched upon as well.
Not quite myriapods, and yet... - Onychophora
As usual we are happy to violate conventional taxonomy and welcome the Onychophora people to our midst (and after all: the myriapods themselves might be a paraphyletic group!). This time there will be three lectures on velvet worms.
Posters
A lecture is just one way of communicating one's observations. Please do not miss the occasion to study the posters which, like the lectures, cover very many aspects of myriapodology. As a small exercise, you can try to fit the posters into the eight abovementioned themes.
Abstracts
Abstracts of lectures and posters are provided in a separate booklet, arranged alphabetically after the first author's surname.
Beyond the themes
Some of the activities of the congress are not of a strict myriapodological nature. This is true of the keynote lecture of Tuesday evening where Jonathan Coddington will speak about a very important subject in today's debate on biodiversity: How can we find out how many species of living organisms there are on Earth?
This is also true of Wednesday's excursion. Although
some may indeed pick up a myriapod or two, the main purpose of
the excursion is to provide you with a day's rest and at the same
time to show a few aspects of Danish nature and culture: the beautiful
forest Jägerspris Nordskov with the almost two thousand years
old "Kongeegen" (Royal Oak), the no less beautiful Frederiksborg
Castle, and finally Frilandsmuseet (The Open Air Museum) with
a selection of old Danish houses placed in a wide open landscape.
The welcome party the light supper served after
the keynote lecture, and the concluding congress dinner
in the Tivoli Gardens are also, on the surface, non ™myriapodological
actvities. But please use these occasions, as well as coffee and
lunch breaks, to establish new contacts and cultivate old ones.
Myriapodology will benefit from this, just as significantly I
think, as from the more formal sessions.
To put it all briefly: Have a nice congress!
On behalf of the organisers
Henrik Enghoff
SUNDAY 28 JULY
1600-1900: Registration at the August Krogh Institute. (Registration also possible during following days)
Welcome party at the Zoological Museum
MONDAY 29 JULY
900-915: Opening of the congress
915-930: Welcome by Professor Niels Peder Kristensen, Department of Entomology, Zoological Museum
First theme: Ancestry and relationships
Chairman: J.-P. Mauriès
930-1000: INVITED LECTURE: Jeffrey W. Shultz & J.C. Regier: Moleclar phylogeny: methodology and application to myriapods.
1000-1030: Coffee break
1030-1100: Wolfgang Dohle: Are the insects more closely related to crustaceans than to the myriapods?
1100-1120: Gero Hilken: On the tracheal systems in Chilopoda: A comparison under phylogenetical aspects.
'1120-1140: Carol-Constantin Prunescu: Contributions to the understanding of the chilopods' evolution.
1140-1200: Joe Hannibal: Pleurojulid millipedes (Diplopoda) from Mazon Creek, Illinois (Carboniferous, USA) and the architecture of the pleurojulid ring structure.(#`
1200-1230: Congress photo at the Niels Bohr monument on the campus lawn.
1230-1400: Lunch. Please remember your ticket
Second theme: Being an arthropod
Chairman: J. Rosenberg
1400-1430: INVITED LECTURE: Michaelis Averof: A genetic view
of arthropod
morphology.
1430-1500: M. Louise Smith & Michael Akam: Hox genes, the evolution of arthropod bodyplans and myriapods.
1500-1530: Coffee break
Chairman: J.-J. Geoffroy
1530-1600: Charles E. Cook & Michael Akam: Hox genes in pauropods and symphylans: tools for the study of phylogeny and body plans.
1600-1630: D. Berto, G. Fusco & Alessandro Minelli: Segmental units and shape control in centipedes.
1630-1645: Bozidar P.M. Curcic & Slobodan Makarov: Postembryonic development of Bulgarosoma lazarevensis Ceuca (Anthroleucosomatidae, Diplopoda) from Yugoslavia.
TUESDAY 30 JULY
Third theme: Order in the diversity
Chairman: J.-F. David
900-920: Jean-Paul Mauriès: Is the statement of the family Atopogestidae (Spirostreptida, Odontopygidae) founded on a case of teratology or a periodomorphotic stage?
920-945: Jörg Spelda: The genus Pyrgocyphosoma Verhoeff, 1910 (Diplopoda, Chordeumatida, Craspedosomatidae): New aspects on systematics, distribution and ecology.
945-1005: Renö Hoess, Adolf Scholl & Mathias Lörtscher: Die Glomeris-Taxa hexasticha Brandt und intermedia Latzel: Arten oder Unterarten? Allozym-Data (Diplopoda, Glomerida, Glomeridae).
1005-1035: Coffee break
Chairman: R.D. Kime
1035-1055: Arkadiy Schileyko: Scolopenders of Vietnam, with special reference to systematics of the Scolopendromorpha (Chilopoda Pleurostigmophora).
1055-1115: Carol-Constantin Prunescu: The structure and the evolution of the genital system in Scolopendromorpha.
1115-1135: Kubra Bano & J.B. Murthy: A comparative account of the gonopods of some paradoxosomatid millipedes.
Fourth theme: Myriapods on Earth
Chairman: A. Minelli
1135-1205: INVITED LECTURE. Sergei I. Golovatch: On the main traits of millipede distribution and faunogenesis in Eurasia.
1205-1220: Bozidar P.M. Curcic & Slobodan Makarov: Diversification and biogeographical features of diplopods (Diplopoda, Myriapoda) in Serbia, Yugoslavia.
1220-1350: Lunch. Please remember your ticket.
Chairwoman: H. Read
1350-1410: Zoltan Korsós: The millipede fauna of the Dráva Region, southern Hungary (Diplopoda).
1410-1430: Kubra Bano & J.B.Murthy: Distribution of the diplopods of the families Harpagophoridae and Paradoxosomatidae in Karnataka State of South India.
1430-1450: INVITED LECTURE. Luis A. Pereira, Donatella Foddai & Alessandro Minelli: Zoogeographical aspects of Neotropical Geophilomorpha.
1450-1510: Heinz-Christian Fründ, Birgit Balkenhol & Birgit Ruszkowski: Chilopoda in forest habitat-islands in North-West Westphalia.
1510-1540: Coffee break
Fifth theme (1): Myriapods in their environments
Chairman: D. Bourdanné
1540-1600: Karel Tajovsky: Distribution of the millipedes (Diplopoda) along an altitudinal gradient in three mountain regions (Czech Republic and Slovak Republic).
1600-1620: R. Desmond Kime: Year-round pitfall trapping of millipedes in Belgium (Diplopoda).
1900-: KEYNOTE LECTURE: Jonathan Coddington: Towards a theory
of Inventory: Estimating species richness.
Royal Danish Academy of Sciences and Letters, H. C. Andersens
Boulevard 35, DK-1552 Copenhagen V. After the lecture a light
supper will be served by the Academy.
Please remember your ticket!
WEDNESDAY 31 AUGUST
900-1800: Excursion to Jøgerspris Nordskov, Frederiksborg
Slot and Frilandsmuseet.
Please remember your ticket. Meeting place: main entrance of the
August Krogh Institute.
THURSDAY 1 AUGUST
Fifth theme (2): Myriapods in their environments
Chairman: U. Scheller
900-930: Daniel Bourdanné: Influence de l'anthropisation sur le peuplement en diplopodes de la région d'Abidjan, Côte d'Ivoire.
930-1000: Tarombera Mwabvu: Millipedes in small-scale farming systems in Zimbabwe: abundance and diversity (Diplopoda, Spirostreptida).
1000-1030: Coffee break
Chairwoman: M. Nguyen Duy-Jacquemin
1030-1100: Dieter Mahsberg: New species - key species: the tropical Pelmatojulus tigrinus Hoffman & Mahsberg, 1996 (Diplopoda, Spirobolida, Pachybolidae).
1100-1130: Somnath Bhakat: Seasonal changes in the distribution of an Indian julid millipede Trigoniulus lumbricinus (Diplopoda, Julida, Trigoniulidae).
1130-1150: Joachim Adis, Ulf Scheller, J.W. de Morais, C. Rochus & J.M.G. Rodrigues: Adaptations of Amazonian Symphyla (Myriapoda) from non-flooded upland forests to innundation forests.
1150-1210: Frank-Thorsten Krell, T. Schmitt & F. Krämer: Scarab beetles (Coleoptera: Scarabaeoidea) specialized on diplopod carcasses (Myriapoda: Diplopoda).
1210-1340: Lunch. Please remember your ticket.
Chairwoman: K. Voigtländer
1340-1400: Peter Bailey: Decline of an invading millipede: A review of theories for Ommatoiulus moreleti in South Australia.
1400-1420: Jolanta Wytwer: Diplopoda communities of inundated ash-alder forests in Puszcza Bialowieska (Poland).
1420-1440: Kubra Bano & J.B. Murthy: Humic fractions from the excreta of two harpagophorid millipedes.
1440-1500: Mark Maraun & Stefan Scheu: Are faecal pellets of millipedes hot spots of microbial activity?
1500-1530: Coffee break
Chairman: K. Ishii
1530-1600: Jean-Jacques Geoffroy: The millipedes of a temperate forest in France: organization and spatio-temporal changes of the community.
1600-1630: Jean-François David & Guy Vannier: First overview of the cold-hardiness of west European millipedes (Diplopoda).
FRIDAY 2 AUGUST
Sixth theme: "A centipede was happy quite"
Chairman: J. G.E. Lewis
900-930: Bruce D. Anderson & Robert J. Full: The biomechanics of metachronal gaits in the centipede, Scolopendra heros.
930-945: Jeffrey W. Shultz, Bruce D. Anderson & B.C. Jayne: Kinematic and electromyographic analysis of axial movement during terrestrial locomotion in the centipede Scolopendra heros (Chilopoda: Scolopendromorpha).
945-1015: Joachim Adis, Gianfranco Caoduro, Benjamin Messner & Henrik Enghoff: On the semiaquatic behaviour of Serradium semiaquaticum, a troglobiotic millipede from northern Italy (Polydesmidae, Diplopoda).
1015-1045: Coffee break
Seventh theme: Myriapods magnified
Chairman: W. Dohle
1045-1115: Monique Nguyen Duy-Jacquemin: Fine structure and possible functions of antennal sensilla in Polyxenus lagurus (Myriapoda, Diplopoda, Penicillata).
1115-1145: Hartmut Greven, Jörg Rosenberg & Inge Latka: The specialized cuticle of the coxal organs of Lithobius forficatus (Chilopoda, Lithobiomorpha): Application of lectins and demonstration of chitin.
1145-1205: Jörg Rosenberg, Evelyn Krüger & Werner Peters: Intense receptor-mediated endocytosis in nephrocytes of myriapods.
1205-1335: Lunch. Please remember your ticket
Eigth theme: Not quite myriapods, and yet...
Chairman: J. Hannibal
1335-1355: Hilke Ruhberg: Ooperipatellus: a unique genus within the oviparous Peripatopsidae (Onychophora).
1355-1415: Claudia Brockmann: Some observations on Ooperipatellus decoratus (Peripatopsidae, Onychophora), an oviparous onychophoran.
1415-1445: Muriel Walker & S.S. Campiglia: The developing
embryo and cyclic changes
in the uterus of a Neotropical onychophoran.
1445-1515: Coffee break
CIM SESSIONS
1515-1600: Business meeting of the Centre International de Myriapodologie.
1600-ca. 1700: Plenary session of the Centre International de Myriapodologie.
1700: Formal closing of the congress.
1930: Congress dinner
Restaurant Påfuglen, Tivoli Gardens. Please remember your
dinner ticket and your ticket to the Tivoli Gardens.
POSTERS
Grzegorz Gacek & Grzegorz Kania: Evidence of prophenoloxidase in the hemolymph of four species of millipedes from Poland.
O. Gealekman, Y. Tichomirova & M.R. Warburg: Life history of a iulid millipede in a Mediterranean pine forest.
Jean-Jacques Geoffroy: The French Millipede Survey: tools and advances.
G. Giribet, A. Serra, S. Carranza, M. Riutori, J. Baguña & C. Ribera: Preliminary internal phylogeny of the Chilopoda (Arthropoda, Myriapoda): a combined approach of complete 18S rDNA and partial 28S rDNA sequences.
Michelle Hamer: A preliminary assessment of southern African millipede (Diplopoda) diversity, distribution and conservation.
Kiyoshi Ishii: Comparative biological study of the penicillate diplopods in Japan.
Grzegorz Kania & Wojciech Rzeski: The phagocytic activity of hemocytes of Ommatoiulus sabulosus.
Ivan Kos: Use of dispersion indexes in ecological research of centipedes (Chilopoda).
Robert Mesibov: Myriapod recording in Tasmania, Australia.
Elena Valentinovna Mikhaljova: Fauna and some ecological particularities of the millipedes (Diplopoda) of the Russian Southern Far East.
J.W. de Morais, J. Adis, E. Berti-Filho, L.A.E. Pereira, A. Minelli & F. Barbieri: On abundance, phenology and natural history of Geophilomorpha (Chilopoda, Myriapoda) from a mixedwater inundation forest in Central Amazonia.
P. Prunescu: The genital system in Lamyctes anderis (Henicopidae, Lithobiomorpha, Chilopoda).
A. Serra, C. Miquel, E. Mateos & C. Vicente: Study of soil Julida communities (Myriapoda, Diplopoda) in Mediterranean forest.
Shernice Soobramoney & Sue L. Marinier: The stridulatory apparatus of two species of pill millipede Sphaerotherium punctulatum and S. giganteum from the coastal forests of Kwazulu-Natal, South Africa.
Willi Xylander & O. Bagisch: The prophenoloxidase of Rhaphidostreptus virgator (Diplopoda Spirostreptidae) is mainly located in the granular hemocytes.
Marzio Zapparoli: Centipedes of a wasteland urban area in Rome, Italy (Chilopoda).
Matthias Zerm: Distribution and phenology of Lamyctes fulvicornis
and other lithobiomorph
centipedes (Chilopoda, Lithobiomorpha) found in floodplains in
the Lower Oder National
Park (Brandenburg, Germany.`
Introduction: Myriapods and myriapodology
by
HENRIK ENGHOFF
Although somewhere between 15,000 and 20,000 species of myriapods
have been described, although several of these are among the most
conspicuous terrestrial invertebrates, and although every child
knows a "millipede", the study of myriapods - myriapodology
- has always been conducted by only a small group of scientists.
In addition to not being very popular as scientific study objects,
myriapods enjoy the dubious distinction of perhaps not representing
a monophyletic group: Some morphological characters indicate the
progoneates: symphylans, pauropods and millipedes (Diplopoda),
to be the sistergroup of the insects, leaving the centipedes (Chilopoda)
outside (Dohle 1980, Kraus & Kraus 1996). Other lines of morphological
evidence, however, suggest monophyly of myriapods, see, e.g. Ax
(1984) and Boudreaux (1979). For instance, the moveable anterior
tentorial apodemes which are unique to myriapods (Manton 1977)
may be a strong synapomorphy for them, but need to be further
evaluated from a phylogenetic point of view. Also in favour of
myriapod monophyly, some molecular phylogenetic studies indicate
centipedes and millipedes to be closer related to each other than
to any other arthropods (Wheeler et al. 1993, Friedrich &
Tautz 1995). Very recently, monophyly of centipedes + symphylans
+ millipedes has been supported by a molecular study (Regier &
Shultz 1997). Pauropods have so far not been included in published
molecular phylogenies.
Even if the myriapods therefore may constitute a monophyletic
group and thus may possess a "genealogical identity",
the position of such a group on the tree of life is debatable.
The commonly held belief has been that myriapods constitute the
sister-group of the insects (Boudraux 1979, Wheeler et al. 1993,
among many others) but other possibilities have been supported
by some recent studies. In this volume, and elsewhere, Dohle (1998)
presents evidence for a sister-group relationship between insects
and crustaceans, thus pushing the myriapods downwards on the arthropod
tree. Several molecular studies tend to support this arrangement,
e.g., Friedrich & Tautz (1995). At least one molecular phylogenetic
study even shows centipedes (other myriapods were not included)
as sister-group to all other arthropods, plus onychophorans (Ballard
et al. 1992, but see Wägele & Stanjek 1995). Evidently,
the study of myriapod relationships is a fruitful field of research,
as is true of many other sub-disciplines of myriapodology (see,
e.g., the papers in the book before you).
Whatever their phylogenetic status - monophyletic or not - myriapods
do form a phenetically relatively homogeneous group: generally,
they are long arthropods with a large number of similar legs.
This myriapodous concept has been usurped by the recently discovered
remipedian crustaceans (Yager 1981) which have even been suggested
as the sistergroup of myriapods and insects (Moura & Christoffersen
1996). Perhaps remipedologists should be invited to future congresses
of myriapodology? This would not be the first adoption by myriapodology
of a non-myriapod group of long, many-legged animals. The study
of onychophorans has traditionally been regarded as lying within
the scope of myriapodology although onychophorans are now believed
to form the sister-group of all arthropods, plus tardigrades (Nielsen
1995).
Since 1968, myriapodologists (including onychophorologists) have
met with about three years' interval, under the auspices of Centre
International de Myriapodologie (C.I.M.). Nearly all of these
congresses have resulted a proceedings volume in which papers
and posters presented at the congress have been published (ninth
congress: Geoffroy et al. 1996; eighth congress: Meyer et al.
1992; earlier congresses: see Meyer et al. 1992: xii). The present
volume contains papers presented at the Tenth International Congress
of Myriapodology which was held in Copenhagen 29 July - 2 August,
1996, sponsored by the C.I.M., the Zoological Museum (University
of Copenhagen), the Danish Natural Science Research Council, the
Royal Danish Academy of Sciences and Letters, the Danish International
Development Agency and Dr. Bøje Benzons Støttefond.
The congress in Copenhagen was attended by 66 myriapodologists
(not counting associates) representing 28 countries in all major
zoogeograpical regions. The scope of the presentations was correspondingly
wide. Those congress papers and posters which are published here
have all been through a standard refereeing procedure which in
some cases has resulted in significant changes from the original
congress contribution. Taken together, the papers advertise that
myriapodology, although an avocation of the selected few, is very
much a living branch of zoology.
Thanks to Wolfgang Dohle, Claus Nielsen, Jørgen Olesen
and Jeffrey Shultz for useful hints.
REFERENCES
Ax, P. 1984. Das phylogenetische System. Systematisierung der
lebenden Natur aufgrund ihrer Phylogenese. Gustav Fischer Verlag,
Stuttgart etc.
Ballard, J.W.O., Olsen, G.J., Faith, D.P., Odgers, W.A., Rowell,
D.M. & Atkinson, P.W. 1992. Evidence from 12S ribosomal RNA
sequences that Onychophora are modified arthropods. - Science
258: 1345-1348.
Boudreaux, H.B. 1979. Arthropod phylogeny with special reference
to insects. John Wiley & sons, New York etc.
Dohle, W. 1980. Sind die Myriapoden eine monophyletische Gruppe?
Eine Diskussion der Verwandschatfsbeziehungen der Antennaten.
- Abh. naturwiss. Ver. Hamburg 23: 45-104.
Dohle, W. 1998. Are the insects more closely related to the crustaceans
than to the myriapods? - Ent. scand. Suppl. 51: 7-16.
Friedrich, M. & Tautz, D. 1995. Ribosomal DNA phylogeny of
the major extant arthropod classes and the evolution of myriapods.
- Nature 376: 165-167.
Geoffroy, J.-J., Mauriès, J.-P. & Nguyen Duy-Jacquemin,
M. (eds.) 1996. Acta Myriapodologica. - Mém. Mus. natn.
Hist. nat. Paris 169: 1-682.
Kraus, O. & Kraus, M. 1996. On myriapod/insect interrelationships.
- Mém. Mus. natn. Hist. nat. 169: 283-290.
Manton, S.M. 1977. The Arthropoda, habits, functional morphology
and evolution. Clarendon Press, Oxford.
Meyer, E., Thaler, K. & Schedl, W. (eds.) 1992. Advances in
myriapodology. Proceedings of the 8th International Congress of
Myriapodology. Ber. Naturw.- Med. Ver. Innsbruck Suppl. 10: i-xiii
+ 1-465.
Moura, G. & Christoffersen, M.L. 1996. The system of the mandibulate
arthropods: Tracheata and Remipedia as sister groups, "Crustacea"
non©monophyletic. - J. Comp. Biol. 1: 95-113.
Nielsen, C. 1995. Animal evolution. interrelationships of the
living phyla. Oxford University Press, Oxford.
Regier, J.C. & Shultz, J.W. 1997. Molecular phylogeny of the
major arthropod groups indicates polyphyly of crustaceans and
a new hypothesis for the origin of hexapods. - Mol. Biol. Evol.
14: 902-913.
Wägele, J.W. & Stanjek, G. 1995. Arthropod phylogeny
inferred from partial 12SrRNA revisited: monophyly of the Tracheata
depends on sequence alignment. - J. zool. Syst. Evol. Res. 33:
75-80.
Wheeler, W.C., Cartwright, P. & Hayashi, C.V. 1993. Arthropod
phylogeny: a combined approach. - Cladistics 9: 1-39.
Yager, J. 1981. Remipedia, a new class of Crustacea from a marine
cave in the Bahamas. - J. Crust. Biol. 1: 328-333.
H. Enghoff, Zoological Museum, University of Copenhagen, Universitetsparken
15, DK©2100 Copenhagen Ø, Denmark. E-mail henghoff@zmuc.ku.dk.
MANY-LEGGED ANIMALS - A collection of papers on Myriapoda and Onychophora
(Proceedings of the Tenth International Congress of Myriapodology,
Copenhagen, 29 July - 2 August, 1996)
HENRIK ENGHOFF (Ed.)
Entomologica scandinavica Supplement 51 (1997)
CONTENTS
The papers are arranged systematically, i.e., classwise. Three papers are on general myriapodology, 12 are on Chilopoda, 24 on Diplopoda, one on Symphyla and one on Onychophora. Within the two larger groups, the papers are arranged, as far as possible, according to the keyword sequence: Phylogeny - systematics - structure - function - distribution - habitats - habits.
Myriapods general
Enghoff, H.: Introduction: myriapods and myriapodology. p. 5
Dohle, W.: Are the insects more closely related to the crustaceans than to the myriapods? p. 7
Rosenberg, J., Krüger, E. & Peters, W.: Intense receptor-mediated endocytosis in nephrocytes of Myriapoda. p. 17
Centipedes, Chilopoda
Shultz, J.W. & Regier, J.C.: Progress toward a molecular phylogeny of the centipede orders (Chilopoda). p. 25
Schileyko, A.A. & Pavlinov, I. Ja.: A cladistic analysis of the order Scolopendromorpha (Chilopoda). p. 33
Prunescu, C.-C.: The anatomy and evolution of the genital system in Scolopendromorpha (Chilopoda). p. 41
Hilken, G.: Tracheal systems in Chilopoda: a comparison under phylogenetic aspects. p. 49
Berto, D., Fusco, G. & Minelli, A.: Segmental units and shape control in Chilopoda. p. 61
Greven, H., Rosenberg, J. & Latka, I.: Cytochemical notes on the specialized cuticle of the coxal organs in Lithobius forficatus: Demonstration of lectins and chitin (Chilopoda, Lithobiomorpha, Lithobiidae). p. 71
Pereira, L.A., Foddai, D. & Minelli, A.: Zoogeographical aspects of Neotropical Geophilomorpha (Chilopoda). p. 77
Stoev, P.: A check-list of the centipedes of the Balkan peninsula with some taxonomic notes and a complete bibliography (Chilopoda). p. 87
Fründ, H.-C., Balkenhol, B. & Ruszkowski, B.: Chilopoda in forest habitat-islands in north-west Westphalia, Germany. p. 107
Morais, J.W. de, Adis, J., Berti-Filho, E., Pereira, L.A., Minelli, A. & Barbieri, F.: On abundance, phenology and natural history of Geophilomorpha from a mixedwater inundation forest in Central Amazonia (Chilopoda). p. 115
Zapparoli, M.: Centipedes of a wasteland urban area in Rome, Italy (Chilopoda). p. 121
Zerm, M.: Distribution and phenology of Lamyctes fulvicornis Meinert, 1868, and other lithobiomorph centipedes in the floodplain of the Lower Oder Valley, Germany (Chilopoda, Henicopidae, Lithobiidae). p. 125
Diplopoda
Hoess, R., Scholl, A. & Lörtscher, M.: The Glomeris-taxa hexasticha and intermedia species or subspecies? - allozyme data (Diplopoda, Glomerida, Glomeridae). p. 133
Mauriès, J.-P.: Is the family Atopogestidae based on a case of teratology or a periodomorphic stage? (Diplopoda, Spirostreptida, Odontopygoidea). p. 139
Sørensen, L.: A new species of the previously monotypic genus Allocotoproctus from the Uluguru Mountains, Tanzania (Diplopoda, Polydesmida, Oxydesmidae). p. 149
Bano, K. & Murthy, J.B.: Gonopods of some South Indian paradoxosomatid millipedes (Diplopoda, Polydesmida). p. 155
Curcic, B.P.M. & Makarov, S.E.: Postembryonic development of Bulgarosoma lazarevensis - a cave-dwelling millipede from Yugoslavia (Diplopoda, Chordeumatida, Anthroleucosomatidae). p. 163
Nguyen Duy-Jacquemin, M.: Fine structure and possible functions
of antennal sensilla in Polyxenus lagurus (Diplopoda, Penicillata,
Polyxenidae. p. 167
Kania, G. & Rzeski, W.: In vitro phagocytic activity of hemocytes
of Ommatoiulus sabulosus: preliminary observations (Diplopoda,
Julida, Julidae). p. 179
Xylander, W.E.R. & Bogusch, O.: Granular hemocytes as the main location of prophenoloxidase in the millipede Rhapidostreptus virgator (Diplopoda, Spirostreptida, Spirostreptidae). p. 183
Curcic, B.P.M. & Makarov, S.E.: Diversification and biogeographic features of millipedes in Serbia, Yugoslavia (Diplopoda). p. 191
Golovatch, S.I.: On the main traits of millipede distribution and faunogenesis in Eurasia (Diplopoda). p. 199
Hamer, M.L.: A preliminary assessment of the southern African millipede fauna: diversity and conservation (Diplopoda). p. 209
Korsós, Z.: The millipede fauna of the Dráva region, southern Hungary (Diplopoda). p. 219
Tajovsky, K.: Distribution of millipedes along an altitudinal gradient in three mountain regions in the Czech and Slovak Republics (Diplopoda). p. 225
Wytwer, J.: Millipede communities of inundated ash-alder forests in Puszcza Bialowieska, Poland (Diplopoda). p. 235
Bailey, P.T.: Decline of an invading millipede, Ommatoiulus moreleti in South Australia: the need for a better understanding of the mechanism (Diplopoda, Julida, Julidae). p. 241
Bourdanné, D.K.: Influence de l'anthropisation sur le peuplement en Diplopodes de la région d'Abidjan, Côte d'Ivoire (Diplopoda). p. 245
David, J.-F. & Vannier, G.: Cold-hardiness of European millipedes (Diplopoda). p. 251
Ishii, K.: Comparative biological study of the penicillate diplopods in Japan (Diplopoda, Penicillata). p. 257
Kime, R.D.: Year-round pitfall trapping of millipedes in mainly open grassland in Belgium (Diplopoda). p. 263
Mahsberg, D.: Pelmatojulus tigrinus, a key detritivore of a tropical gallery forest, (Diplopoda, Spirobolida, Pachybolidae). p. 269
Krell, F.-T., Schmitt, T. & Linsenmair, K.E.: Diplopod defensive secretions as attractants for necrophagous scarab beetles (Diplopoda - Insecta, Coleoptera, Scarabaeidae). p. 281
Mwabvu, T.: Millipedes in small-scale farming systems in Zimbabwe: abundance and diversity (Diplopoda, Spirostreptida). p. 287
Serra, A., Miquel, C., Mateos, E. & Vicente, C.: Study of a soil Julidae community in Mediterranean forest (Diplopoda, Julida). p. 291
Adis, J., Caoduro, G., Messner, B. & Enghoff, H.: On the semiaquatic behaviour of a new troglobitic millipede from northern Italy (Diplopoda, Polydesmida, Polydesmidae). p. 301
The smaller groups: Symphyla and Onychophora
Adis, J., Scheller, U., Morais, J.W. de, Rochus, C. & Rodrigues, J.M.G.: Symphyla from Amazonian non-flooded upland forests and their adaptations to inundation forests. p. 307
Brockmann, C., Mesibov, R. & Ruhberg, H.: Observations on Ooperipatellus decoratus, an oviparous onychophoran from Tasmania (Onychophora, Peripatopsidae). p. 319